"This foundation [for sociobiologists concerned with cooperation] includes Hamilton’s reformulation of natural selection as a force that maximizes “inclusive fitness” rather than individual fitness, George Williams’ expository evisceration of old-school group selection, Robert Trivers’ explanation of cooperation between unrelated individuals with the theory of reciprocity, and John Maynard Smith’s application of game theory to the study of animal behavior. Sadly, too few of us include George Price in this august company."